The roller on the transfer unit at the K toner end appears to have started to disintegrate and is very deformed. It has also pierced the belt which runs across it, meaning the entire transfer unit will need replacing at a cost of about £272 for parts (+vat).
The faulty part can be removed by: removing all toner cartridges, clicking out the 4 vertical pieces on the right hand side (looking from front to back) and undoing the 2 screws holding the transfer unit.
We may instead go for a new printer!
Once again, thanks for all your help.